For almost 40 years, Dr. Anthony S. Fauci has held two jobs. As director of the National Institute of Allergy and Infectious Diseases, he has run one of the country’s premier research institutions. But he has also been an adviser to seven presidents, from Ronald Reagan to, now, Joseph R. Biden Jr., called upon whenever a health crisis looms to brief the administration, address the World Health Organization, testify before Congress or meet with the news media.

For Dr. Fauci, 80, the past year has stood out like no other. As the coronavirus ravaged the country, Dr. Fauci’s calm counsel and commitment to hard facts endeared him to millions of Americans. But he also became a villain to millions of others. Trump supporters chanted “Fire Fauci,” and the president mused openly about doing so. He was accused of inventing the virus and of being part of a secret cabal with Bill Gates and George Soros to profit from vaccines. His family received death threats. On Jan. 21, appearing in his first press briefing under the Biden administration, Dr. Fauci described the “liberating feeling” of once again being able to “get up here and talk about what you know — what the evidence, what the science is — and know that’s it, let the science speak.”

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